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Publishing Director: Linda Belfus Acquisitions Editor: William Schmitt Managing Editor: Rebecca Gruliow Publishing Services Manager: Tina Rebane Project Manager: Mary Anne Folcher Design Manager: Steven Stave Marketing Manager: John Gore

Cover illustration is a detail from Opus 1972 by Virgil Cantini, Ph.D., with permission of the artist and Mansfield State College, Mansfield, Pennsylvania.

Chapter opener credits: Chapter 43, modified from © Getty Images 21000058038;Chapter 44, modified from © Getty Images 21000044598;Chapter 84, modified from © Corbis.

Printed in China

Last digit is the print number: 987654321

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ELSEVIER Sabre Foundation

My Family

For their abundant support, for their patience and understanding, and for their love

Arthur C. Guyton For his imaginative and innovative research For his dedication to education For showing us the excitement and joy of physiology And for serving as an inspirational role model

Arthur C. Guyton, M.D. 1919-2003

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Essentials of Human Physiology

Essentials of Human Physiology

This ebook provides an introductory explanation of the workings of the human body, with an effort to draw connections between the body systems and explain their interdependencies. A framework for the book is homeostasis and how the body maintains balance within each system. This is intended as a first introduction to physiology for a college-level course.

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