Neuronal Effector Mechanisms That Decrease or Increase Body Temperature

When the hypothalamic temperature centers detect that the body temperature is either too high or too low, they institute appropriate temperature-decreasing or temperature-increasing procedures. The reader is probably familiar with most of these from personal experience, but special features are the following.

Temperature-Decreasing Mechanisms When the Body Is Too Hot

The temperature control system uses three important mechanisms to reduce body heat when the body temperature becomes too great:

1. Vasodilation of skin blood vessels. In almost all areas of the body, the skin blood vessels become intensely dilated. This is caused by inhibition of the sympathetic centers in the posterior hypothalamus that cause vasoconstriction. Full vasodilation can increase the rate of heat transfer to the skin as much as eightfold.

2. Sweating. The effect of increased body temperature to cause sweating is demonstrated by the blue curve in Figure 73-7, which shows a sharp increase in the rate of evaporative heat loss resulting from sweating when the body core temperature rises above the critical level of 37°C (98.6°F). An additional 1°C increase in body temperature causes enough sweating to remove 10 times the basal rate of body heat production.

3. Decrease in heat production. The mechanisms that cause excess heat production, such as shivering and chemical thermogenesis, are strongly inhibited.

Effect of hypothalamic temperature on evaporative heat loss from the body and on heat production caused primarily by muscle activity and shivering. This figure demonstrates the extremely critical temperature level at which increased heat loss begins and heat production reaches a minimum stable level.

Temperature-Increasing Mechanisms When the Body Is Too Cold

When the body is too cold, the temperature control system institutes exactly opposite procedures. They are:

1. Skin vasoconstriction throughout the body. This is caused by stimulation of the posterior hypothalamic sympathetic centers.

2. Piloerection. Piloerection means hairs "standing on end." Sympathetic stimulation causes the arrector pili muscles attached to the hair follicles to contract, which brings the hairs to an upright stance. This is not important in human beings, but in lower animals, upright projection of the hairs allows them to entrap a thick layer of "insulator air" next to the skin, so that transfer of heat to the surroundings is greatly depressed.

3. Increase in thermogenesis (heat production). Heat production by the metabolic systems is increased by promoting shivering, sympathetic excitation of heat production, and thyroxine secretion. These methods of increasing heat require additional explanation, which follows.

Hypothalamic Stimulation of Shivering. Located in the dorsomedial portion of the posterior hypothalamus near the wall of the third ventricle is an area called the primary motor center for shivering. This area is normally inhibited by signals from the heat center in the anterior hypothalamic-preoptic area but is excited by cold signals from the skin and spinal cord. Therefore, as shown by the sudden increase in "heat production" (see the red curve in Figure 73-7), this center becomes activated when the body temperature falls even a fraction of a degree below a critical temperature level. It then transmits signals that cause shivering through bilateral tracts down the brain stem, into the lateral columns of the spinal cord, and finally to the anterior motor neurons. These signals are non-rhythmical and do not cause the actual muscle shaking. Instead, they increase the tone of the skeletal muscles throughout the body by facilitating the activity of the anterior motor neurons. When the tone rises above a certain critical level, shivering begins. This probably results from feedback oscillation of the muscle spindle stretch reflex mechanism, which is discussed in Chapter 54. During maximum shivering, body heat production can rise to four to five times normal.

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Essentials of Human Physiology

Essentials of Human Physiology

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Responses

  • Monte
    What effector mechanisms for increasing and decreasing body temperature?
    2 years ago
  • Alaric
    How do effectors decrease body temperature?
    1 year ago

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